Worldwide.

Plants terrestrial, on rock, or epiphytic. Roots emerging near origin, or growing through cortex and emergent some distance from origin. Horizontal stems present or absent, mainly protostelic, in some species becoming actino- or plectostelic, on substrate surface or subterranean, or forming stolons. Upright shoots simple or branched, usually conspicuously leafy at least at base; abscising gemmae formed by reduced lateral shoots. Lateral shoots present or absent, simple or branched, branching pattern dichotomous and sometimes pseudomonopodial; leaves uniform or dimorphic or trimorphic. Upright and lateral shoots round or flat in cross section; leaves on subterranean parts flat, appressed, nonphotosynthetic, and scalelike; leaves on aerial parts appressed, ascending, or spreading, with 1 central unbranched vein, needlelike to lanceolate to ovate, remote to dense and imbricate, with or without basal and/or mucilage canals. Strobili sessile or stalked, upright, nodding, or pendent. Sporangia solitary, adaxial near leaf base or axillary; subtending leaves (sporophylls) unmodified and photosynthetic to much modified, nonphotosynthetic, reduced, and aggregated in strobili; sporangia reniform to globose, thick-walled with hundreds of spores, outer walls variously modified. Spores all 1 kind, trilete, thick-walled, surfaces pitted to small-grooved, rugulate, or reticulate. Gametophytes subterranean and nonphotosynthetic or surficial and photosynthetic.
(Flora of North America)