|
|
Taxon
|
Location
|
Latitude
|
Longitude
|
Date
|
Method
|
Ecology
|
Host plant
|
Observer
|
Specimens
|
Types
|
Museum
|
Reference
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515603
|
34.807206
|
14.10.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
5
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.511822
|
34.816459
|
14.10.2015
|
pitfall trap
|
foothill psammophyte grassland
|
|
Sumarokov A.M.
|
16
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.513768
|
34.824957
|
14.10.2015
|
pitfall trap
|
foothill psammophyte grassland
|
|
Sumarokov A.M.
|
2
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.521728
|
34.814861
|
04.10.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
2
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.514912
|
34.823421
|
07.09.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.522301
|
34.822321
|
07.09.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.512204
|
34.805406
|
07.09.2015
|
pitfall trap
|
uphill psammophyte grassland
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.518414
|
34.81726
|
07.09.2015
|
pitfall trap
|
pine forest
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.520519
|
34.804117
|
19.08.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.514912
|
34.823421
|
19.08.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515603
|
34.807206
|
19.08.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
2
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.518414
|
34.81726
|
19.08.2015
|
pitfall trap
|
pine forest
|
|
Sumarokov A.M.
|
2
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.520519
|
34.804117
|
08.08.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.514912
|
34.823421
|
08.08.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515603
|
34.807206
|
08.08.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
18
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515264
|
34.805543
|
08.08.2015
|
pitfall trap
|
pine forest
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.518414
|
34.81726
|
08.08.2015
|
pitfall trap
|
pine forest
|
|
Sumarokov A.M.
|
2
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.51628
|
34.822336
|
29.07.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
5
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.514912
|
34.823421
|
29.07.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
6
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515603
|
34.807206
|
29.07.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
28
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.524741
|
34.814737
|
29.07.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.518414
|
34.81726
|
29.07.2015
|
pitfall trap
|
pine forest
|
|
Sumarokov A.M.
|
8
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.521728
|
34.814861
|
20.07.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515541
|
34.824253
|
20.07.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
5
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.520059
|
34.816194
|
20.07.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
6
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.511069
|
34.815773
|
20.07.2015
|
pitfall trap
|
Acer tatarica shrubland
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.521728
|
34.814861
|
29.06.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515541
|
34.824253
|
29.06.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.522301
|
34.822321
|
29.06.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515541
|
34.824253
|
17.06.2015
|
pitfall trap
|
marsh
|
|
Sumarokov A.M.
|
1
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.520519
|
34.804117
|
17.06.2015
|
pitfall trap
|
oak forest
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
|
Dniprovsko-Orelskyi Nature Reserve
|
48.515603
|
34.807206
|
17.06.2015
|
pitfall trap
|
meadow
|
|
Sumarokov A.M.
|
3
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Pavlohrad raion_
|
Pavlohrad
|
48.49042
|
35.944383
|
01.05.2008
|
human observation
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Pavlohrad raion_
|
Pavlohrad
|
48.49042
|
35.944383
|
01.05.2008
|
human observation
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Pavlohrad raion_
|
Pavlohrad
|
48.49042
|
35.944383
|
01.05.2008
|
human observation
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Kyivska oblast
|
Bila Tserkva raion_
|
|
49.747398
|
30.159691
|
16.07.2022 17:29:12
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Dnipro raion_
|
|
48.466347
|
35.005582
|
12.10.2021 15:28:56
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Dnipro raion_
|
|
48.466347
|
35.005582
|
12.10.2021 15:29:29
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Khersonska oblast
|
Kherson raion_
|
|
46.653833
|
32.558239
|
24.06.2021
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Mykolaivska oblast
|
Mykolaiv City
|
Ингульский р-н, ул. Космонавтов
|
46.945451
|
32.058377
|
27.05.2021 17:47:22
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Kyivska oblast
|
Bila Tserkva raion_
|
|
49.80281
|
30.13476
|
09.04.2020 14:08:43
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Kyivska oblast
|
Bila Tserkva raion_
|
|
49.80281
|
30.13476
|
09.04.2020 14:17:36
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Dnipro raion_
|
|
48.439916
|
35.010198
|
25.05.2019 14:29:52
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Dnipro raion_
|
|
48.439916
|
35.010198
|
25.05.2019 14:30:21
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Zakarpatska oblast
|
Berehove raion_
|
смт.Королево
|
48.151833
|
23.137822
|
27.06.2018
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Cherkaska oblast
|
Mankivka raion
|
|
48.978044
|
30.250889
|
19.06.2018 11:23:23
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Pavlohrad raion_
|
Pavlohrad
|
48.49042
|
35.944383
|
01.05.2008 15:27:24
|
|
|
|
|
|
|
|
|
|
|
det. А.Гуж (A.Gorz)
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Pavlohrad raion_
|
Pavlohrad
|
48.49042
|
35.944383
|
01.05.2008 15:28:29
|
|
|
|
|
|
|
|
|
|
|
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
|
Ukraine
|
Dnipropetrovska oblast
|
Pavlohrad raion_
|
Pavlohrad
|
48.49042
|
35.944383
|
01.05.2008 15:28:44
|
|
|
|
|
|
|
|
|
