Distribution
Ukraine

Native to: Alabama, Alberta, Altay, Austria, Baltic States, Belarus, Belgium, British Columbia, Central European Rus, Connecticut, Czechoslovakia, Delaware, Denmark, District of Columbia, East European Russia, Finland, France, Georgia, Germany, Great Britain, Hungary, Idaho, Illinois, Indiana, Iowa, Italy, Krasnoyarsk, Maine, Manitoba, Maryland, Masachusettes, Michigan, Minnesota, Missouri, Montana, Nebraska, Netherlands, New Brunswick, New Hampshire, New Jersey, New York, Newfoundland, North Carolina, North Dakota, North European Russi, Northwest European R, Norway, Nova Scotia, Ohio, Ontario, Pennsylvania, Poland, Prince Edward I., Québec, Rhode I., Romania, Saskatchewan, Sweden, Switzerland, Tennessee, Ukraine, Vermont, Virginia, Washington, West Siberia, West Virginia, Wisconsin, Yugoslavia.

Leaves dimorphic, 35-70 × 8-12 cm; fertile leaves dying back in winter; sterile leaves several, small, green through winter, forming "rosette". Petiole 1/4-1/3 length of leaf, scaly at least at base; scales scattered, tan. Blade green, narrowly lanceolate or with parallel sides, pinnate-pinnatifid, not glandular. Pinnae of fertile leaves twisted out of plane of blade and perpendicular to it, deltate; basal pinnae deltate, somewhat reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal; pinnule margins distantly serrate, with spiny teeth. Sori midway between midvein and margin of segments. Indusia lacking glands. 2 n = 164.
Swamps, swampy woods, or open shrubby wetlands; 0-1200 m.