Distribution
Czech Republic, Hungary, Ukraine

Native to: Alabama, Albania, Alberta, Altay, Arkansas, Austria, Baltic States, Belarus, Belgium, British Columbia, Bulgaria, Buryatiya, Central European Rus, Chita, Connecticut, Corse, Czechoslovakia, Delaware, Denmark, District of Columbia, East European Russia, Finland, France, Georgia, Germany, Great Britain, Hungary, Idaho, Illinois, Indiana, Iowa, Ireland, Irkutsk, Italy, Kentucky, Krasnoyarsk, Krym, Labrador, Maine, Manitoba, Maryland, Masachusettes, Michigan, Minnesota, Missouri, Montana, Nebraska, Netherlands, New Brunswick, New Hampshire, New Jersey, New York, Newfoundland, North Carolina, North Dakota, North European Russi, Northwest European R, Northwest Territorie, Norway, Nova Scotia, Nunavut, Ohio, Ontario, Oregon, Pennsylvania, Poland, Prince Edward I., Québec, Rhode I., Romania, Saskatchewan, South Carolina, South Dakota, Spain, Sweden, Switzerland, Tennessee, Transcaucasus, Turkey, Ukraine, Vermont, Virginia, Washington, West Siberia, West Virginia, Wisconsin, Xinjiang, Yugoslavia, Yukon.

Leaves monomorphic, dying in winter, 15-75 × 10-30 cm. Petiole 1/4-1/3 length of leaf, scaly at least at base; scales scattered, tan. Blade light green, ovate-lanceolate, 2-3-pinnate-pinnatifid, herbaceous, not glandular. Pinnae ± in plane of blade, lance-oblong; basal pinnae lanceolate-deltate, slightly reduced, basal pinnules usually longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. Sori midway between midvein and margin of segments. Indusia lacking glands. 2 n = 164.
Swampy woods, moist wooded slopes, stream banks, and conifer plantations; 0-1200 m.